Zonation-relevant references (loosely grouped)
Primarily methodological
Oldest basic Zonation references
Moilanen, A. 2007. Landscape Zonation, benefit functions and target-based planning. Unifying reserve selection strategies. Biological Conservation, 134: 571-579.
Moilanen, A., Franco, A.M.A., Early, R., Fox, R., Wintle, B., and C.D. Thomas. 2005. Prioritising multiple use landscapes for conservation: methods for large multi species planning problems. Proc. R. Soc. Lond. B Biol. Sci., 272: 1885-1891.
Three more recent pieces about generally relevant methods
Moilanen, A., and Arponen A. 2011. Administrative regions in conservation: balancing local priorities with regional to global preferences in spatial planning. Biological Conservation, 144: 1719-1725.
Moilanen, A., B.J. Anderson, F. Eigenbrod, A. Heinemeyer, D. B. Roy, S. Gillings, P. R. Armsworth, K. J. Gaston, and C.D. Thomas. 2011. Balancing alternative land uses in conservation prioritization. Ecological Applications, 21: 1419-1426.
Lehtomäki, J. and A. Moilanen. 2013. Methods and workflow for spatial conservation prioritization using Zonation. Environmental Modelling & Software, 47: 128-137.
Application in impact avoidance and/or biodiversity offsetting
Kareksela, S., Moilanen, A., Ristaniemi, O., Välivara, R. and J.S. Kotiaho. 2017. Exposing ecological and economic costs of the research implementation gap and compromises in decision making. Conservation Biology 32, 9-17.
Moilanen, A. 2013. Planning impact avoidance and biodiversity offsetting using software for spatial conservation prioritization. Wildlife Research, 40: 153-162.
Kareksela, S., Moilanen, A., Tuominen, S. and Kotiaho J.S. 2013. Use of Inverse spatial conservation prioritization to avoid biodiversity loss outside protected areas. Conservation Biology, 27: 1294–1303.
Community or environment -level analysis methods
Leathwick, J.R., Moilanen A., Ferrier S. and Julian K.. 2010. Community-level conservation prioritization and its application to riverine ecosystems. Biological Conservation, 143, 984-991.
Moilanen, A., J.R. Leathwick, and J. M. Quinn. 2011. Spatial prioritization of conservation management. Conservation letters, 4, 383-393. Biological Conservation, 160, 234-241.
Sharafi, S.M. , Moilanen A., White, M., and Burgman M.A. 2012. Integrating environmental gap analysis with spatial conservation prioritization: A case study from Victoria, Australia. Journal of Environmental Management, 112, 240-251.
Arponen, A.; Heikkinen, R.K.; Paloniemi, R.; Pöyry, J.; Similä, J. & Kuussaari, M. 2013. Improving conservation planning for semi-natural grasslands: Integrating connectivity into agri-environment schemes.
Harlio, A.; Kuussari, M.; Heikkinen, R.K. & Arponen, A. 2019. Incorporating landscape heterogeneity into multi-objective spatial planning improves biodiversity conservation of semi-natural grasslands. Journal of Nature Conservation, 49, 37-44.
BQP connectivity method; distribution smoothing
Moilanen, A. and B. A. Wintle. 2007. The boundary quality penalty a quantitative method for approximating species responses to fragmentation in reserve selection. Conservation Biology, 21: 355-364.
Why use Zonation in the first place instead of target-based systematic conservation planning
Moilanen, A., and Arponen A. 2011a. Setting conservation targets under budgetary constraints. Biological Conservation, 144: 650-653.
Di Minin, E. and A. Moilanen. 2012. Empirical evidence for reduced protection levels across biodiversity features from target-based conservation planning. Biological Conservation, 153: 187-191.
Laitila, J. and A. Moilanen. 2013. Use of many low-level conservation targets reduces high-level conservation performance. Ecological Modelling, in press.
Interactions and interaction connectivity between features
Rayfield, B., Moilanen, A. and Fortin M.-J. 2009. Incorporating consumer-resource spatial interactions in reserve design. Ecological Modelling, 220: 725-733.
Lehtomäki, J., Tomppo, E., Kuokkanen, P. Hanski, I., and A. Moilanen. 2009. Planning of forest conservation areas using high-resolution GIS data and software for spatial conservation prioritization. Forest Ecology and Management, 258: 2439-2449.
Freshwater (tree topology) connectivity
Moilanen, A., Leathwick, J.R., and J. Elith. 2008. A method for freshwater conservation prioritization. Freshwater Biology, 53: 577-592.
Uncertainty analysis
Moilanen, A. and B.A. Wintle. 2006. Uncertainty analysis favours selection of spatially aggregated reserve structures. Biological Conservation, 129: 427-434.
Moilanen, A., B.A. Wintle., J. Elith and M. Burgman. 2006a. Uncertainty analysis for regional-scale reserve selection. Conservation Biology 20: 1688-1697.
Edge artefacts in national-level planning
Moilanen, A., Anderson, B.J., Arponen, A., Pouzols, F.M., and C.D. Thomas. 2013. Edge artefacts and lost performance in national versus continental conservation priority areas. Diversity and Distributions 19: 171–183.
Applications with useful analysis setups
Lehtomäki, J., Kusumoto, B., Shiono, T., Tanaka, T., Kubota, Y. & Moilanen, A. 2018. Spatial conservation prioritization for the East Asian islands: A balanced representation of multitaxon biogeography in a protected area network. Diversity and Distributions 2018: 1-16. https://doi.org/10.1111/ddi.12869
Virtanen, E. A., Viitasalo, M., Lappalainen, J. & Moilanen, A. 2018. Evaluation, Gap Analysis, and Potential Expansion of the Finnish Marine Protected Area Network.Frontiers in Marine Science 5: 1-19. https://doi.org/10.3389/fmars.2018.00402
Pouzols, F.M. , Toivonen, T,. Di Minin, E., Kukkala, A., Kullberg, P., Kuusterä,J, Lehtomäki, J., Tenkanen, H., Verburg, P.H. & Moilanen, A. (2014) Global protected area expansion is compromised by projected land-use and parochialism. Nature 516: 383–386.
Design of reserve networks and surrogacy
Kareksela, S., Aapala, K., Alanen, A., Haapalehto, T., Kotiaho, J. S., Lehtomäki, J., Leikola, N., Mikkonen, N., Moilanen, A., Nieminen, E., Tuominen, S. & Virkkala, R. 2020. Combining spatial prioritization and expert knowledge facilitates effectiveness of large-scale mire protection process in Finland. Biological Conservation 241: 8 s. https://doi.org/10.1016/j.biocon.2019.108324
Kremen, C., A. Cameron, A. Moilanen, S. Phillips, C. D. Thomas, H. Beentje, J. Dransfeld, B. L. Fisher, F. Glaw, T. Good, G. Harper, R.J. Hijmans, D. C. Lees, E. Louis Jr., R. A. Nussbaum, A. Razafimpahanana, C. Raxworthy, G. Schatz, M. Vences, D. R.Vieites, P. C. Wright , M. L.Zjhra. 2008. Aligning conservation priorities across taxa in Madagascar, a biodiversity hotspot, with high-resolution planning tools. Science, 320: 222-226.
Franco, A.M.A., Anderson, B., Roy, D.B., Gillings, S., Fox,R., Moilanen, A., and C.D. Thomas. 2009. Surrogacy and persistence in reserve selection: landscape prioritisation for multiple taxa in Britain. Journal of Applied Ecology, 46: 82-91.
Carbon and biodiversity
Thomas, C.D., Anderson, B.J., Moilanen, A., Eigenbrod, F., Heinemeyer, A., Quaife T., Roy, D.B., Gillings, S., Armsworth, P.R., and K. J. Gaston. 2013. Reconciling biodiversity and carbon conservation. Ecology Letters, 16-S1: 39-47.
Evaluation of proposed conservation areas, good example
Leathwick, J. R., Moilanen, A., Francis, M., Elith, J., Taylor, P. Julian, K. and T. Hastie. 2008. Novel methods for the design and evaluation of marine protected areas in offshore waters. Conservation Letters, 1: 91-102.
Habitat restoration planning
Thomson, J.R., Moilanen, A., McNally, R., and P. Vesk. 2009. A quantitative method for prioritizing landscape revegetation. Ecological Applications, 19: 817-828.
Applications in the context of climate change
Carroll, C., Moilanen, A., and J. Dunk. 2010. Designing multi-species reserve networks for resilience to climate change: priority areas for spotted owl and localized endemics in the pacific North-West USA. Global Change Biology, 16: 891-904.
Kujala, H, Moilanen, A., Araújo, M.B., and M. Cabeza. 2013. Conservation planning with uncertain climate change projections. PLoS One 8: e53315, doi:10.1371/journal.pone.0053315.
Expansion of conservation areas accounting for connectivity
Lehtomäki, J., Tomppo, E., Kuokkanen, P. Hanski, I., and A. Moilanen. 2009. Planning of forest conservation areas using high-resolution GIS data and software for spatial conservation prioritization. Forest Ecology and Management, 258: 2439-2449.
Urban environments
Gordon, A., Simondson, D., White, M., Moilanen, A., and Bekessy, S.A. 2009. Integrating conservation planning and land-use planning in urban landscapes. Landscape and Urban Planning, 91: 183-194.
Bekessy, S.A., White, M., Gordon, A., Moilanen, A., Mccarthy, M.A.,and B.A. Wintle. 2012. Transparent planning for biodiversity and development in the urban fringed. Landscape and Urban Planning, 108: 140-149.
Single-species planning
Sirkiä, S., Lehtomäki, J., Lindén,H., Tomppo, E. and A Moilanen. 2013. Spatial conservation prioritization of capercaillie (Tetrao urogallus) lekking landscapes in South-Central Finland. Wildlife Biology, in press.
Analysis of national Natura 2000 network
Mikkonen, N. and A. Moilanen. 2013. Identification of top priority areas and management landscapes from a national Natura 2000 network. Environmental Science & Policy, 27: 11-20.
Application to genetic data
Taberlet, P., Gugerli, F., Zimmermann, N.E., Englisch, T., Tribsch, A., Holderegger, R., Alvarez, N., Niklfeld, H., Mirek, Z., Moilanen, A., et al. (54 authors). 2012. Genetic diversity in widespread species is not congruent with species richness in alpine plant communities. Ecology Letters 15: 1439-1448.
Ecotourism, balancing multiple land uses
Di Minin, E., MacMillan, D.C., Goodman, P.S., Escott, B., Slotow, R., and A. Moilanen. 2013. Businesses and conservation planning in a biological diversity hotspot. Cons. Biol., 27: 808-820.
Z-relevant Book Chapters
Moilanen, A., Kujala, H. and J. Leathwick. 2009. The Zonation framework and software for conservation prioritization. Pp. 196-210 in Moilanen, Wilson and Possingham (Eds.), Spatial Conservation Prioritization, Oxford University Press.
Moilanen, A. and I. Ball. 2009. Heuristic and approximate optimization methods for spatial conservation prioritization. Pp. 58-69 in Moilanen, Wilson and Possingham (Eds.), Spatial Conservation Prioritization, Oxford University Press.
The manuals
Newest
Di Minin, E., V. Veach, F.M. Pouzols, J. Lehtomäki and A. Moilanen. 2014. A quick introduction to Zonation, 32 pp. ISBN: 978-952-10-9921-2 (paperback)
Moilanen, A., F.M. Pouzols, L. Meller, V. Veach, A. Arponen, J. Leppänen, and H. Kujala. 2014. Zonation spatial conservation planning methods and software V4, User manual, 290 pp. ISBN 978-952-10-9919-9 (paperback)
Old versions
Moilanen, A., L. Meller, J. Leppänen, H. Kujala and A. Arponen. 2011. Zonation spatial conservation planning framework and software v3.0, User manual, ~300 pp. http://www.helsinki.fi/bioscience/consplan/software/Zonation/downloads.html
Moilanen, A. and H. Kujala. 2008. Zonation spatial conservation planning framework and software v. 2.0, User manual, 136 pp. www.helsinki.fi/bioscience/consplan.
Moilanen, A. and H. Kujala. 2006. Zonation spatial conservation planning framework and software v. 1.0, User manual, 126 pp. Edita, Helsinki, Finland.
Other relevant bits of methods not specific to Zonation
Review of the concepts of spatial prioritization
Kukkala, A. and A. Moilanen. 2013. The core concepts of spatial prioritization in systematic conservation planning. Biological Reviews, 88: 443-464.
Book about spatial prioritization
Moilanen, A., K. A. Wilson, and H.P. Possingham, editors. 2009. Spatial Conservation Prioritization: Quantitative Methods and Computational Tools. Oxford University Press.
What data has typically been used in SCP
Kullberg, P. and A. Moilanen. 2014. Spatial biodiversity analyses and information needs of the Convention on Biological Diversity. Natureza & Conservacao, 12: 3-10.
Mathematical classification of conservation prioritization problems
Moilanen, A., Possingham, H.P., and S. Polasky. 2009. A mathematical classification of conservation prioritisation problems. Pp. 28-42 in Moilanen, Wilson and Possingham (Eds.), Spatial Conservation Prioritization, Oxford University Press.
Community level analysis – why it works or not
Arponen, A., Moilanen, A., and S. Ferrier. 2008. A successful community-level strategy for conservation prioritization. Journal of Applied Ecology, 45: 1436-1445.
Replacement cost
Cabeza, M. and A. Moilanen. 2006. Replacement cost: a useful measure of site value for conservation planning. Biological Conservation 132: 336-342.
Moilanen, A., A. Arponen, J. Stokland and M. Cabeza. 2009. Assessing replacement cost of conservation areas: how does habitat loss influence priorities? Biological Conservation, 142: 575-585.
Generalized complementarity – dependence and independence between planning units
Moilanen, A. 2008b. Generalized Complementarity and Mapping of the Concepts of Systematic Conservation Planning. Conservation Biology, 22: 1655-1658.
The ecological model and optimization: what is it that matters?
Moilanen, A. 2008a. Two paths to a suboptimal solution: once more about optimality in reserve selection. Biological Conservation 141: 1919-1923.
Info-gap reserve selection
Moilanen, A., M. Runge, J. Elith, A. Tyre, Y. Carmel, E. Fegraus, B. Wintle, M. Burgman and Y. Ben-Haim. 2006b. Planning for robust reserve networks using uncertainty analysis. Ecological Modelling, 119: 115-124.
Focus on consequences of conservation actions instead of pattern-based spatial planning
Pouzols, F.M., Burgman, M.A., and A. Moilanen. 2012. Methods for allocation of habitat management, maintenance, restoration and offsetting, when conservation actions have uncertain consequences. Biological Conservation, 153: 41-50.
Pouzols, F.M. and A. Moilanen. 2013. RobOff: software for analysis of alternative land-use options and conservation actions. Methods in Ecology and Evolution, 4: 426-432.